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Publications and Reports
Publications and Reports

​​​​​​​​​​​​​Wildlife Research ​​​​​​​​​Annual Reports

Wildlife Research Report​​​​s


Baigas, P. E., J. R. Squires, L E. Olson, J. S. Ivan, E. K. Roberts. 2017. Using environmental features to model highway crossing behavior of Canada lynx in the Southern Rocky Mountains. Landscape and Urban Planning 157:200–213;

Ivan, J. S., and T. M. Shenk. 2016. Winter diet and hunting success of Canada lynx in Colorado. Journal of Wildlife Management 80:1049-1058; DOI: 10.1002/jwmg.21101​

Buderman, F.E., Hooten M.B., Ivan, J.S., Shenk, T.M. 2016. A functional model for characterizing long distance movement behavior. Methods in Ecology and Evolution 7:264–273; DOI: 10.1111/2041-210X.12465

Ivan, J. S. In Press. Canada lynx (Lynx Canadensis). Pages 000–000 in J. L. E. Cartron and J. K. Frey, editors. The Wild Carnivores of New Mexico. University of New Mexico Press, Albuquerque, USA.

Trainor, A. M., O. J. Schmitz, J. S. Ivan, and T. M. Shenk. 2014. Enhancing species distribution modeling by characterizing predator–prey interactions. Ecological Applications 24:204–216: DOI: 10.1890/13-0336.1.

Mountain Lion

Moss, W. E., M. W. Alldredge, and J. N. Pauli. 2016. Quantifying risk and resource use of a large carnivore in an expanding urban–wildland interface. Journal of Applied Ecology 53:371-378; doi: 10.1111/1365-2664.12563

Moss, W. E., M. W. Alldredge, K. A. Logan, and J. N. Pauli. 2016. Human expansion precipitates niche expansion for an opportunistic apex predator (Puma concolor). Scientific Reports 6:39639; doi: 10.1038/srep39639​

Blecha, K. A. and M. W. Alldredge. 2015. Improvements on GPS location cluster analysis for the prediction of large carnivore feeding activities: ground-truth detection probability and inclusion of activity sensor measures. PLoS ONE 10(9):e0138915.doic10.1371/journal.pone.0138915.

Lee, J., S.N. Bevins, L. Serieys, W. Vickers, K.P. Logan, M.W. Alldredge, E. Boydston, L. Lyren, R. McBride, M. Roelke‐Parker, J. Pecon‐Slattery, J. Troyer, S. Riley, W.M. Boyce, K. Crooks, and S. VandeWoude. 2014. The evolution of puma lentivirus in bobcats (Lynx rufus) and mountain lions (Puma concolor) in North America. Journal of Virology 88(14): 7727-7737; doi: 10.1128/JVI.00473-14.

Black Bear

Kirby, R., M. W. Alldredge, and J. N. Pauli. 2017. Environmental, not individual, factors drive markers of biological aging in black bears. Evolutionary Ecology 31: 571-581; 

Kirby, R., M.W. Alldredge, and J.N. Pauli.  2016. The diet of black bears tracks the human footprint across a heterogeneous landscape.  Biological Conservation 200:51-59;

Wolfe, L. L., H. E. Johnson, M. C. Fisher, W. R. Lance, D. K. Smith, and M. W. Miller. 2016. Chemical immobilization in American black bears using a combination of nalbuphine, medetomidine, and azaperone. Ursus 27:1–4; DOI: 10.2192/URSUS-D-15-00018.1

Alldredge, M.W., D.P. Walsh, L.L. Sweanor, R.B. Davies, and A. Trujillo. 2015. Evaluation of translocation of black bears involved in bear-human conflicts in south-central Colorado. Wildlife Society Bulletin; DOI: 10.1002/wsb.526.

Johnson, H. E., S. W. Breck, S. Baruch-Mordo, D.L. Lewis, C.W. Lackey, K.R. Wilson, J. Broderick, J. Mao, and J.P. Beckmann. 2015. Shifting perceptions of risk and reward: dynamic selection for human development by black bears in the western United States. Biological Conservation 187:164-172;

Wolfe, L. L., H. E. Johnson, M. C. Fisher, M. A. Sirochman, B. Kraft, and M. W. Miller. 2014. Use of acepromazine and medetomidine in combination for sedation and handling of Rocky Mountain elk (Cervus elaphus) and black bears (Ursus americanus). Journal of Wildlife Diseases 50:979-981; DOI: 10.7589/2014-02-052.


Johnson, H.E., J.W. Fischer, M. Hammond, P.D. Dorsey, C. Anderson, and K.C. VerCauteren. 2014. Evaluat​​​​ion of techniques to reduce deer and elk damage to agricultural crops​. Wildlife Society Bulletin 38:358-365.

Brodie, J., H. Johnson, M. Mitchell, P. Zager, K. Proffitt, M. Hebblewhite, M. Kauffman, B. Johnson, J. Bissonette, C. Bishop, J. Gude, K. Hersey, M. Hurley, P. Lukacs, S. McCorquodale, E. McIntire, J. Nowak, D. Smith, and P.J. White. 2013. Relative influence of human harvest, carnivores, and weather on adult female elk survival across western North America​. ​Journal of Applied Ecology 50:295-305.


Nadeau, M. S., N. J. DeCesare, D. G. Brimeyer, E. J. Bergman, R. B. Harris, K. R. Hersey, K. K. Huebner, P. E. Mathews, and T. P. Thomas. 2017. Status and trends of moose populations and hunting opportunity in the western United States. Alces 52:99-112.​

Mule Deer

Bergman, E. J., C. R. Anderson, Jr., C. J. Bishop, A. A. Holland, and J. M. Northrup. 2017. Variation in ungulate body fat: Individual versus temporal effects. Journal of Wildlife Management; DOI: 10.1002/jwmg.21334.

Stonehouse, K. F., C. R. Anderson Jr., M. E. Peterson, and D. R. Collins. 2016. Approaches to field investigations of cause-specific mortality in mule deer (Odocoileus hemionus). Colorado Parks and Wildlife Technical Report No. 48, First Edition, 317 W. Prospect Rd., Ft. Collins, CO USA. DOW-R-T-48-16, ISSN 0084-8883.

Northrup, J. M., C. R. Anderson, Jr., M. B. Hooten, and G. Wittemyer. 2016. Movement reveals scale dependence in habitat selection of a large ungulate. Ecological Applications 26:2746-2757

Northrup, J. M., C. R. Anderson, Jr., and G. Wittemyer. 2016. Environmental dynamics and anthropogenic development alter philopatry and space-use in a North American cervid. Diversity and Distributions 22: 547-557; DOI: 10.1111/ddi.12417

Johnson, H. E., J. R . Sushinsky, A. Holland, E. J. Bergman , T. Balzer, J. Garner and S. E. Reed. 2016. Increases in residential and energy development are associated with reductions in recruitment for a large ungulate. Global Change Biology, doi: 10.1111/gcb.13385

Bergman, E. J., P. F. Doherty Jr., G. C. White, and A. A. Holland. 2015. Density dependence in mule deer: a review. Wildlife Biology 21:18-29; doi: 10.2981/wlb.00012

Bergman, E. J., P. F. Doherty Jr., G. C. White, and D. J Freddy. 2015. Habitat and herbivore density: response of mule deer to habitat management. Journal of Wildlife Management 79(1):60–68; 2015; DOI: 10.1002/jwmg.801

Northrup, J. M., C. R. Anderson, Jr., and G. Wittemyer. 2015. Quantifying spatial habitat loss from hydrocarbon development through assessing habitat selection patterns of mule deer. Global Change Biology, doi: 10.1111/gcb.13037

Searle, K. R., M. B. Rice, C. R. Anderson, C. Bishop and N. T. Hobbs. 2015. Asynchronous vegetation phenology enhances winter body condition of a large mobile herbivore. Oecologia 179:377-391; DOI 10.1007/s00442-015-3348-9

Anderson, C. R., Jr., and C. J. Bishop. 2014. Migration patterns of adult female mule deer in response to energy development. Pages 46-50 in R. A. Coon & M. C. Dunfee, editors. Transactions of the 79th North American Wildlife & Natural Resources Conference. Wildlife Management Institute, Gardners, PA, USA. ISSN 0078-1355.

Bergman, E. J., P. F. Doherty, Jr., C. J. Bishop, L. L. Wolfe, and B. A. Banulis. 2014. Herbivore body condition response in altered environments: mule deer and habitat management. PLoS ONE 9(9): e106374. doi:10.1371/journal.pone.0106374

Bergman, E.J., C.J. Bishop, D.J. Freddy, G.C. White and P.F. Doherty. ​2014. Habitat management influence overwinter survival of mule deer fawns in Colorado. Journal of Wildlife Management 78(3):448–455; DOI: 10.1002/jwmg.683

Freeman, E. D., R. T. Larsen, M. E. Peterson, C. R. Anderson, Jr., K. R. Hersey, B. R. McMillan. 2014. Effects of male‐biased harvest on mule deer: implications for rates of pregnancy, synchrony, and timing of parturition​. ​Wildlife Society Bulletin; DOI: 10.1002/wsb.450.

Lendrum, P. E., C. R. Anderson, Jr., R. A. Long, J. G. Kie, and R. T. Bowyer. 2014. Habitat selection by mule deer during migration: effects of landscape structure and natural-gas development. Pages 233-262 in M. McBroom, editor. The Effects of Hydraulic Fracturing on the Environment. Apple Academic Press Inc., Waretown, NJ, USA.​

Lendrum, P. E., C. R. Anderson, Jr., K. L. Monteith, J. A. Jenks, R. T. Bowyer. 2014. Movements of a rapidly migrating ungulate: effects of forage quality. Mammalian Biology​ 79:369-375;

​Lynch, E., J.M. Northrup, M.F. McKenna, C.R. Anderson, Jr., L, Angeloni, and G. Wittemyer. 2014. Landscape and anthrogenic features influencing the use of auditory vigilance by mule deer. Behavioral Ecology; doi:10.1093/beheco/aru158.

Northrup, J. M., A. B. A. Shafer, C. R. Anderson, Jr., D. W. Coltman, and G. Wittemyer. 2014. Fine-scaled genetic correlates to condition and migration in a wild cervid. Evolutionary Applications ISSN 1752-4571.

Northrup, J. M., C. R. Anderson, Jr., and G. Wittemyer. 2014. Effects of helicopter capture and handling on movement behavior of mule deer​. Journal of Wildlife Management; DOI: 10.1002/jwmg.705.

Bergman, E.J. 2013. Evaluation of winter range habitat treatments on overwinter survival, density, and body condition of mule deer​. Dissertation, Colorado State University, Fort Collins, Colorado, USA, 137pp.

Bergman, E.J., C.J. Bishop, D.J. Freddy, G.C. White and P.F. Doherty. 2014. Habitat management influence overwinter survival of mule deer fawns in Colorado​. Journal of Wildlife Management 78(3):448–455.

Other Publications:

Hooten, M. B., F. E. Buderman, B. M. Brost, E. M. Hanks, and J. S. Ivan. 2016. Hierarchical animal movement m​​odels for population-level inference​. Environmetrics 27:322-333.

Carver, S, S. N. Bevins, M. R. Lappin, E. E. Boydston, L. M. Lyren, M. Alldredge, K. A. Logan, L. L. Sweanor, S. P. D. Riley, L. E. K. Serieys, R. N. Fisher, T. W. Vickers, W. Boyce, R. McBride, M. C. Cunningham, M. Jennings, J. Lewis, T. Lunn, K. R. Crooks, S.VandeWoude.2016. Pathogen exposure varies widely among sympatric populations of wild and domestic felids across the United States. Ecological Applications 26:367-381 doi: 10.1890/15-0445.1

Ivan, J. S., and E. S. Newkirk. 2015. CPW Photo Warehouse: a custom database to facilitate archiving, identifying, summarizing, and managing photo data collected from camera traps. Methods in Ecology and Evolution. doi: 10.1111/2041-210X.12503.​

Hanks, E.M., M.B. Hooten, and M. Alldredge. 2015. Continuous-time discrete-space models for animal movement. Annals of Applied Statistics, 9: 145-165.

Lewis, J.S., K. A. Logan, M.W. Alldredge, L.L. Bailey, S. VandeWoude, K.R. Crooks. 2015. The effects of urbanization on population density, occupancy, and detection probability of wild felids. Ecological Applications 25: 1880-1895.

Gerber, B. D., J. S. Ivan, and K. P. Burnham. 2014. Estimating the abundance of rare and elusive carnivores from photographic - sampling data when the population size is very small. Population Ecology 56:463-479; DOI: 10.1007/s10144-014-0431-8.

Hooten, M. B., E. M. Hanks, D. S. Johnson, and M. W. Alldredge. 2014. Temporal variation and scale in movement‐based resource selection functions. Statistical Methodology 17:82‐98

Ivan, J. S., G. C. White, and T. M. Shenk. 2014. Density and demography of snowshoe hares in central Colorado. Journal of Wildlife Management 78:580-594; DOI: 10.1002/jwmg.695

Mills. L. S., and H. E. Johnson. 2013. Wildlife Population Dynamics, pages 84‐109 in Wildlife management: contemporary principles and practices. John Hopkins University Press.

Hooten, M. B., E. M. Hanks, D. S. Johnson, and M. W. Alldredge. 2013. Reconciling resource utilization and resource selection functions. Journal of Animal Ecology 82: 1146‐1154; doi: 10.1111/1365-2656.12080

Northrup, J. M., M. B. Hooten, C. R. Anderson, Jr., and G. Wittemyer. 2013. Practical guidance on characterizing availability in resource selection functions under a use - availability design. Ecology 94(7):1456‐1463; DOI: 10.1890/12-1688.1.

Ellis, M. M., J. S. Ivan, and M. K. Schwartz. 2013. Spatially explicit power analyses for occupancy‐based monitoring of wolverine in the U.S. Rocky Mountains. Conservation Biology 28:52–62; 10.1111/cobi.12139