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Jake Ivan
Jake Ivan

​​​Wildlife Researcher, Species of Conservation Concern


  • Ph.D., Fish, Wildlife and Conservation Biology — Colorado State University, 2011
  • M.S., Wildlife Biology — University of Montana, 2000
  • B.S., Wildlife Science — Purdue University, 1997

Current or Recent Positions

  • Wildlife Researcher — Colorado Parks and Wildlife, 2010​ – Present
  • Refuge Biologist — U.S. Fish and Wildlife Service, 2003-2005
  • Piping Plover Recovery Biologist — The Nature Conservancy, 2000-2003

Areas of Interest, Expertise

My research focuses on estimating wildlife population parameters and forest carnivore ecology, conservation and monitoring. 

Current or Recent Research Projects

  • Impacts of pine and spruce beetle outbreaks on breeding birds and mammals in Colorado.

  • Identification of movement corridors for lynx in Colorado.

  • Effects of winter recreation on the movements and resource use of Canada Lynx in Colorado.

Select Publications

  • Baigas, P. E., J. R. Squires, L. E. Olsen, J. S. Ivan, and E. K. Roberts. In Press. Using environmental features to model highway crossing behavior of Canada lynx in the Southern Rocky Mountains. Landscape and Urban Planning 00:000-000.

  • Hooten, M. B., F. E. Buderman, B. M. Brost, E. M. Hanks, and J. S. Ivan. In Press. Hierarchical animal movement models for population-level inference. Environmetrics. 00:000-000.

  • Ivan, J. S., and T. M. Shenk. 2016. Winter diet and hunting success of Canada lynx in Colorado. Journal of Wildlife Management DOI:10.1002/jwmg.21101

  • Ivan, J. S., and E. S. Newkirk. 2016. Cpw Photo Warehouse: a custom database to facilitate archiving, identifying, summarizing, and managing photo data collected from camera traps. Methods in Ecology and Evolution 7:499-504.

  • Buderman F. E., M. B. Hooten, J. S. Ivan, and T. M. Shenk. 2016. A functional model for characterizing long distance movement behavior. Methods in Ecology and Evolution 7:264-273. ​

  • Ellis, M. M., J. S. Ivan, J. M. Tucker, and M. K. Schwartz. 2015. rSPACE: Spatially based power analysis for conservation and ecology. Methods in Ecology and Evolution 6:621-625.​

  • Ivan, J. S. In Press. Canada lynx (Lynx Canadensis). Pages 000–000 in J. L. E. Cartron and J. K. Frey, editors. The Wild Carnivores of New Mexico. University of New Mexico Press, Albuquerque, New Mexico, USA. 000 pp.

  • Ivan, J. S., G. C. White, and T. M. Shenk. 2014. Density and ​demography of snowshoe hares in central Colorado.  Journal of Wildlife Management 78:580–594.

  • Gerber, B. D., J. S. Ivan, and K. P. Burnham. 2014. Estimating the abundance of rare and elusive carnivores from photographic-sampling data when the population size is very small. Population Ecology 56:463-470.

  • Trainor, A. M., O. J. Schmitz, J. S. Ivan, and T. M. Shenk. 2014. Enhancing species distribution modeling by characterizing predator–prey interactions. Ecological Applications 24:204–216.

  • Ellis, M. M., J. S. Ivan, and M. K. Schwartz. 2013. Spatially explicit power analyses for occupancy-based monitoring of wolverine in the U.S. Rocky Mountains. Conservation Biology 28:52–62.

  • Ivan, J. S., G. C. White, and T. M. Shenk. 2013. Using auxiliary telemetry information to estimate animal density from capture-recapture data. Ecology 94(4): 809–816.

  • Ivan, J. S., G. C. White, and T. M. Shenk. 2013. Using simulation to compare methods for estimating density from capture-recapture data. Ecology 94 (4): 817–826.

  • Ivan, J. S. 2013. Density estimation. Pages 20.1-20.19 in E. Cooch and G. White, Editors. Program MARK - 'A Gentle Introduction'. 11th Edition. 983pp. "Program MARK" software overview, book and download.

  • Stromborg, K. L., J. S. Ivan, J. K. Netto, and C. R. Courtney. In press. Survivorship and mortality patterns of cormorants at Spider Island, Wisconsin, 1988-2006. Waterbirds 35:31-39.

  • Ivan, J. S., and R. K. Murphy. 2005. What preys on piping plover eggs and chicks? Wildlife Society Bulletin 33:113-119.